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My name is Chaitanya (first name) Gokhale (last name). I was born in Pune, which is a city in Central Western India (about 200 kms inland from Mumbai). Below is a web version of my curriculum vitae which can also be downloaded as a pdf below.

Awards and Grants

  • Otto Hahn Medal
  • Principal Investigator - Partial Grant from the DFG Priority Programme `Probabilistic Structures in Evolution’ (SPP 1590)
  • BMC Ecology Image Competition: Theoretical Ecology and Models ___

April 2016 -

March 2014 - March 2016

March 2011 - February 2014

May 2008 - March 2011

July 2005 - January 2008

  • Master of Science in Bioinformatics
    Thesis: “Ab initio calculations on the HGPRT active site and the analysis of select mutations” (Score: 187/200)
    Advisor: Dr. Mrinalini Puranik

June 2002 - June 2005

  • Bachelor of Science in Zoology and Biotechnology Fergusson College, Pune University


Editorial Board

  • Journal of Evolutionary Biology
  • Scientific Reports


  • Dynamic Games and Applications
  • Ecology Letters
  • Evolution
  • Journal of Mathematical Biology
  • Journal of The Royal Society Interface
  • Journal of Theoretical Biology
  • Mathematical Biosciences
  • Proceedings of the National Academy of Sciences, USA
  • Proceedings of the Royal Society B: Biological Sciences
  • PLoS Computational Biology
  • PLoS One
  • Theoretical Population Biology


  • Santa Fe Complex Systems Summer School (CSS) applications
  • L’OREAL Austria [Fellowship for Young Female Scientists in Basic Research]

Academic Experience


  1. Czuppon, P. and Gokhale, C. S. (2018). Disentangling eco-evolutionary effects on trait fixation, Theoretical Population Biology
  2. Roemhild , R. et al. (2018). Cellular hysteresis as a principle to maximize the efficacy of antibiotic therapy, PNAS
  3. Daubech et al. (2017). Spatio-temporal control of mutualism in legumes helps spread symbiotic nitrogen fixation, eLife
  4. Schenk, H., Traulsen, A. and Gokhale, C.S. (2017). Chaotic provinces in the kingdom of the Red Queen, Journal of Theoretical Biology
  5. Gokhale, C.S., Traulsen, A. and Joop, G. (2016). Social Dilemma In The External Immune System Of The Red Flour Beetle? It Is A Matter Of Time, Ecology and Evolution
  6. Liu, Y., Gokhale, C.S., Rainey, R.B. and Zhang, X-X. (2017). Unravelling the complexity and redundancy of carbon catabolic repression in the plant growth‐promoting bacterium Pseudomonas fluorescens SBW25, Molecular Microbiology
  7. Bertels, F., Gokhale, C.S. and Traulsen, A. (2016). Discovering complete quasispecies in bacterial genomes, Genetics
  8. Gokhale, C. S. and Hauert, C. (2016). Eco-evolutionary dynamics of social dilemmas, Theoretical Population Biology
  9. Papkou, A., Gokhale, C. S., Traulsen, A. & Schulenburg, H. (2016). Host parasite co-evolution, Why changing population size matters, Zoology
  10. Song, Y., Gokhale, C. S., Papkou, A., Schulenburg, H. & Traulsen, A. (2015). Host-parasite coevolution in populations of constant and variable size. BMC Evolutionary Biology, 15, 212.
  11. Bauer, B. & Gokhale, C.S., Repeatability of evolution on epistatic landscapes. Scientific reports, 5, p.9607. (2015)
  12. "Mathematica Interactive CDF published on Wolfram Demonstrations"
  13. Gokhale, C. S., Reeves R. G. & Reed, F.A. (2014). Dynamics of a combined Medea-Underdominant Population Transformation System. BMC Evolutionary Biology, 14(1), 98
  14. Gokhale, C. S., & Traulsen, A. (2014). Evolutionary Multiplayer Games. Dynamic Games and Applications.
  15. Gokhale, C. S., Papkou, A., Traulsen, A., & Schulenburg, H. (2013). Lotka-Volterra dynamics kills the Red Queen: population size fluctuations and associated stochasticity dramatically change host-parasite coevolution. BMC Evolutionary Biology, 13(1), 254.
  16. "Mathematica Interactive CDF published on Wolfram Demonstrations"
  17. Wu B, Traulsen A, Gokhale CS. Dynamic Properties of Evolutionary Multi-player Games in Finite Populations. Games. Multidisciplinary Digital Publishing Institute; 2013 May 6;4(2):182–99.
  18. Wu B, Gokhale CS, Veelen M, Wang L. Interpretations arising from Wrightian and Malthusian fitness under strong frequency dependent selection. Ecology and Evolution. 2013.
  19. Gokhale CS, Traulsen A. Mutualism and evolutionary multiplayer games: revisiting the Red King. Proceedings of the Royal Society B: Biological Sciences. 2012 Oct 11;279(1747):4611–6.
  20. Han TA, Traulsen A, Gokhale CS. On equilibrium properties of evolutionary multi-player games with random payoff matrices. Theoretical Population Biology. 2012 Jun;81(4):264–72.
  21. Wu B, Gokhale CS, Wang L, Traulsen A. How small are small mutation rates? J. Math. Biol. 2011 May 28;64(5):803–27.
  22. Gokhale CS, Traulsen A. Strategy abundance in evolutionary many-player games with multiple strategies. JTB. 2011 Aug;283(1):180–91.
  23. Altrock PM, Gokhale CS, Traulsen A. Stochastic slowdown in evolutionary processes. Phys. Rev. E. 2010 Jul;82(1 Pt 1):011925.
  24. Gokhale CS, Traulsen A. Evolutionary games in the multiverse. Proceedings of the National Academy of Sciences. 2010 Mar 23;107(12):5500–4.
  25. Bhardwaj A, Mukerji M, Sharma S, Paul J, Gokhale CS, Srivastava AK, et al. MtSNPscore: a combined evidence approach for assessing cumulative impact of mitochondrial variations in disease. BMC Bioinformatics. 2009;10(Suppl 8):S7.
  26. Gokhale CS, Iwasa Y, Nowak MA, Traulsen A. The pace of evolution across fitness valleys. JTB. 2009 Aug;259(3):613–20.