Chaitanya S. Gokhale

Chaitanya S. Gokhale

Theorist. Hiker. Likes it cold.

Insight must precede application

Max Planck

If the paper you are interested in is not freely available then drop me an email and I will send it.

Preprints

  • Lagos-Oviedo, J.J., Rajendra, D and Gokhale, C. S. and Schmitt, T and Frank, E.T., Evolution of first aid and social wound care in an army ant society bioRxiv-2025.09.22.677721
  • Gokhale, C. S. and Traulsen, A. Higher-order equivalence of Lotka-Volterra and replicator dynamics bioRxiv-2025.03.28.645916
  • Krishnan, N., Kun, Á., Gokhale, C. S. and Garay, J. Early eukaryogenesis by host-initiated obligate ectosymbiosis and metabolic inhibition. bioRxiv-2024.09.08.611858
  • Lauenroth, D., and Gokhale, C. S. Assessing the probability and time of weed control failure due to herbicide resistance evolution. bioRxiv-2024.04.26.591266v1
  • Suswaram, D., Yeakel, J.D. and Gokhale, C. S. Parasitoid pressures and silence evolution bioRxiv-12.503800
  • Bargués-Ribera, M., Reeves, R.G., Gokhale, C. S., Eco-evolutionary dynamics of Plasmodium genotypes under mass drug administration, bioRxiv-818039
  • Rabajante, J.F., Anzia, E.L., Gokhale, C. S., On a mechanistic process of macroparasite aggregation, bioRxiv-680041
  • Gokhale, C. S., Wignall, A.E., On the innovation and evolution of predatory tactics, bioRxiv-530238

Book review

  • Gokhale, C. S. and Traulsen, A.T., Rethinking the origins of biological games, reviewing Game Theory in Biology: Concepts and Frontiers by John M. McNamara and Olof Leimar for the International Society for Behavioural Ecology newsletter

Peer-reviewed

Journal articles

         2026

  1. Patil, S., Nöbel, S., & Gokhale, C. S. Phenotypic polymorphism via mate copying, Proc. Natl. Acad. Sci. U.S.A. 123 (9) e2510849123, https://doi.org/10.1073/pnas.2510849123

  2. Rajendra,D., Gokhale, C. S. Optimizing play for learning risky behaviour. Proc. Roy. Soc. Biol. Sci. 2026; 293 (2065): 20253111. https://doi.org/10.1098/rspb.2025.3111

     2025
  3. Athreya, G. S., Gokhale, C. S. & Verma, P. Antibiotic-mediated interactions underlying microbial diversity. PLoS Computational Biology forthcoming bioRxiv-2023.02.15.528676
  4. Arango, A., Peters, M.K. & Gokhale, C. S. (2025), History, climate, and competition shape ecological specialization in the Emberizoidea radiation. J Avian Biol, 2025: e03506. https://doi.org/10.1002/jav.03506
  5. Athreya, G. S., Czuppon, P. & Gokhale, C. S. The evolution of dependence and cohesion in incipient endosymbioses. The American Naturalist (2025) link
  6. W. Barfuss, J. Flack, Gokhale, C. S., L. Hammond, C. Hilbe, E. Hughes, J.Z. Leibo, T. Lenaerts, N. Leonard, S. Levin, U. Madhushani Sehwag, A. McAvoy, J.M. Meylahn, & F.P. Santos, Collective cooperative intelligence, Proc. Natl. Acad. Sci. U.S.A. 122 (25) e2319948121, pdf
  7. Mikaberidze A, Gokhale, C. S., Bargués-Ribera M, Verma P (2025) The cost of fungicide resistance evolution in multi-field plant epidemics. PLoS Sustain Transform 4(6): e0000178. pdf
  8. Fic M., Bastian F., Miękisz J., and Gokhale, C. S. Compartment model of strategy-dependent time delays in replicator dynamics. Journal of Theoretical Biology, 601, 2025, 112044 pdf arXiv:2409.01116

  9. Nguyen, P. L., & Gokhale, C. S. On multiple infections by parasites with complex life cycles. OIKOS pdf bioRxiv-2023.10.13.562184v1

     2024
  10. Fic, M., Gokhale, C. S. Catalysing cooperation: the power of collective beliefs in structured populations. npj Complexity 1, 6 (2024). pdf

  11. Revathi Venkateswaran, V., Gokhale, C. S., Mangel, M., and Eliassen, S., Effects of time spent in pregnancy or brooding on immunocompetence Ecology & Evolution 14,1 e10764 pdf

     2023
  12. Gokhale, C. S., Rainey, P. B., and Frean, M., Eco-evolutionary logic of mutualisms, Dynamic Games and Applications pdf
  13. Lauenroth, D., and Gokhale, C. S. Theoretical assessment of persistence and adaptation in weeds with complex life cycles, Nature Plants 9, 1267-1279 bioRxiv-12.503772
  14. Gokhale, C. S., and Sharma, N. 2023 Optimising crop rotations via Parrondo’s paradox for sustainable agriculture. Royal Society Open Science 10:221401 Royal Soc. Open Science

  15. Gokhale, C. S., Velasque, M. and Denton, J. A. 2023 Ecological Drivers of Community Cohesion. mSystems, 10.1128/msystems.00929-22 bioRxiv-22022.05.15.491981v1

     2022
  16. Verma, P., Reeves, R. G., Simon, S. Otto, M. and Gokhale, C. S. 2022 The effect of mating complexity on gene drive dynamics. The American Naturalist, 201, 1 bioRxiv-2021.09.16.460618
  17. Gokhale, C. S., Bulbulia, J. & Frean, M. 2022 Collective narratives catalyse cooperation. Humanit Soc Sci Commun 9, 85 socRxiv socRxiv
  18. Gao, Y., Pichugin, Y., Gokhale, C. S. & Traulsen, A. 2022 Evolution of reproductive strategies in incipient multicellularity. J Roy Soc Interface 19, 20210716. bioRxiv-2021.09.13.460035

  19. Traulsen, A., Gokhale, C. S., Shah, S. & Uecker, H. The Covid-19 pandemic: Basic insights from basic mathematical models, NAL Live - Nationale Akademie Leopoldina

     2021
  20. Gokhale, C. S., Giamo, S., & Remigi, P. 2021 Memory shapes microbial populations, PLoS Computational Biology, 17, e1009431 bioRxiv-370106
  21. Verma, P., Reeves, R. G. & Gokhale, C. S. 2021 A common gene drive language eases regulatory process and eco-evolutionary extensions. BMC Ecol Evol 21, 156. bioRxiv-970103
  22. Mafessoni, F., Lachmann, M., Gokhale, C. S., 2021 On the fitness of informative cues in complex environments. Journal of Theoretical Biology, 527, 110819 bioRxiv-066571
  23. Venkateswaran, V. R., Roth, O. & Gokhale, C. S. 2021 Consequences of combining sex‐specific traits. Evolution doi:10.1111/evo.14204. bioRxiv-version-892810

  24. Park, H.J.$ , Gokhale, C. S.$ , Bertels, F. ($joint first authors), 2021. How sequence populations persist inside bacterial genomes. Genetics doi:10.1093/genetics/iyab027 bioRxiv-version-170514

    2020
  25. Denton, J.A., Gokhale, C. S., (joint first authors) 2020. Synthetic symbiosis under environmental disturbances. mSystems, 5 (3) bioRxiv version-395426

  26. Bargués-Ribera, M., Gokhale, C. S., 2020. Eco-evolutionary agriculture: host-pathogen dynamics in crop rotations. PLOS Computational Biology, 16(1): e1007546 bioRxiv-version-402313

    2019
  27. Venkateswaran, V.R., Gokhale, C. S., 2019. Evolutionary dynamics of complex multiple games. Proceedings of the Royal Society B: Biological Sciences,286(1905), p.20190900 OpenAccess bioRxiv-302265
  28. Denton, J.A., Gokhale, C. S., 2019. Synthetic mutualism and the intervention dilemma. Life - Special Issue “Synthetic Biology: From Living Computers to Terraformation”, 9 (1), 15

  29. Park, H-J., Gokhale, C. S., 2019. Ecological feedback on diffusion dynamics. Journal of the Royal Society Open Science, 6 (2), 181273

     2018
  30. Czuppon, P, Gokhale, C. S., 2018. Disentangling eco-evolutionary effects on trait fixation. Theoretical Population Biology, 124, 93-107.

  31. Roemhild, R., Gokhale, C. S., Dirksen, P., Blake, C., Rosenstiel, P., Traulsen, A., Andersson, D., Schulenburg, H., 2018. Cellular hysteresis as a novel principle to maximize the efficacy of antibiotic therapy, PNAS, 115(39), 9767-9772.

     2017
  32. Daubech, B., Remigi, P., Doin de Moura, G., Marchetti, M., Pouzet, C., Auriac, M-C., Gokhale, C. S., Masson-Biovin C., Capela, D., 2017. Spatio-temporal control of mutualism in legumes helps spread symbiotic nitrogen fixation, eLife, 6, 57.
  33. Schenk, H., Traulsen, A., Gokhale, C. S., 2017. Chaotic provinces in the kingdom of the Red Queen. Journal of Theoretical Biology, 431, pp.1–10.
  34. Liu, Y., Gokhale C.S., Rainey P.B., Zhang, X-X., 2017. Unravelling the complexity and redundancy of carbon catabolic repression in Pseudomonas fluorescens SBW25. Molecular microbiology, 105(4), pp.589–605.
  35. Bertels, F., Gokhale, C. S., Traulsen, A., 2017. Discovering Complete Quasispecies in Bacterial Genomes. Genetics, 206(4), pp.2149–2157.

  36. Gokhale, C. S., Traulsen, A., Joop, G., 2017. Social dilemma in the external immune system of the red flour beetle? It is a matter of time. Ecology & Evolution, 7(17), pp.6758–6765.

    2016
  37. Gokhale, C. S., Hauert, C., 2016. Eco-evolutionary dynamics of social dilemmas. Theoretical Population Biology, 111, 28–42.
  38. Papkou, A., Gokhale, C. S., Traulsen, A., Schulenburg, H., 2016. Host-parasite coevolution: why changing population size matters. Zoology (Jena, Germany), 119(4), pp.330–338. 
    2015
  39. Pichugin, Y., Gokhale, C. S., Garcia, J., Traulsen, A., Rainey, P.B., 2015. Modes of migration and multilevel selection in evolutionary multiplayer games. Journal of Theoretical Biology, 387, 144–153.
  40. Song, Y. Gokhale, C. S., Papkou, A., Schulenburg, H.,Traulsen, A., 2015. Host-parasite coevolution in populations of constant and variable size. BMC Evolutionary Biology, 15(1), p.212.

  41. Bauer, B., Gokhale, C. S., 2015. Repeatability of evolution on epistatic landscapes. Scientific reports, 5, p.9607.

    2014
  42. Gokhale, C. S., Traulsen, A., 2014. Evolutionary Multiplayer Games. Dynamic Games and Applications, 4(4), pp.468–488.

  43. Gokhale, C. S., Reeves, R.G., Reed, F.A., 2014. Dynamics of a combined medea-underdominant population transformation system. BMC Evolutionary Biology, 14(1), 98.

    2013
  44. Gokhale, C. S., Papkou, A., Traulsen, A., Schulenburg, H., 2013. Lotka-Volterra dynamics kills the Red Queen: population size fluctuations and associated stochasticity dramatically change host-parasite coevolution. BMC Evolutionary Biology, 13(1), p.254.
  45. Wu, B., Gokhale C.S., van Veelen, M., Wang, L., Traulsen, A., 2013. Interpretations arising from Wrightian and Malthusian fitness under strong frequency dependent selection. Ecology & Evolution, 3(5), pp.1276–1280.

  46. Wu, B., Traulsen, A., Gokhale, C. S., 2013. Dynamic Properties of Evolutionary Multi-player Games in Finite Populations. Games, 4(2), pp.182–199.

    2012
  47. Gokhale, C. S., Traulsen, A., 2012. Mutualism and evolutionary multiplayer games: revisiting the Red King. Proceedings of the Royal Society B: Biological Sciences, 279(1747), 4611–4616.
  48. Wu, B., Gokhale C.S., Wang, L., Traulsen, A., 2012. How small are small mutation rates? Journal of Mathematical Biology, 64(5), pp.803–827.

  49. Han, T.A., Traulsen, A., Gokhale, C. S., 2012. On equilibrium properties of evolutionary multi-player games with random payoff matrices. Theoretical Population Biology, 81(4), pp.264–272.

    2011

  50. Gokhale, C. S., Traulsen, A., 2011. Strategy abundance in evolutionary many-player games with multiple strategies. Journal of Theoretical Biology, 283, 180–191.

    2010
  51. Altrock, P.M., Gokhale, C. S., Traulsen, A., 2010. Stochastic slowdown in evolutionary processes. Physical Review E, 82(1), 11925.

  52. Gokhale, C. S., Traulsen, A., 2010. Evolutionary games in the multiverse. PNAS, 107(12), 5500–5504.

    2009
  53. Gokhale, C. S., Iwasa, Y., Nowak, M., Traulsen A., 2009. The pace of evolution across fitness valleys. Journal of Theoretical Biology, 259(3), 613–620.
  54. Bhardwaj, A., Mukerji, M., Sharma, S., Paul, J., Gokhale, C. S., Srivastava, A.K., Tiwari, S., 2009. MtSNPscore: a combined evidence approach for assessing cumulative impact of mitochondrial variations in disease., BMC Bioinformatics, 10 Suppl 8, S7.

Book Chapter

        2020

  1. Gokhale, C. S., Park, H-J., Eco-evolutionary Spatial dynamics of Non-linear Social Dilemmas, in Advances in Dynamic Games, eds. David M. Ramsey and Jérôme Renault bioRxiv version-660266